Ecological Archives E096-194-A3

A. Coulon, J. Aben, S. C. F. Palmer, V. M. Stevens, T. Callens, D. Strubbe, L. Lens, E. Matthysen, M. Baguette, and J. M. J. Tavis. 2015. A stochastic movement simulator improves estimates of landscape connectivity. Ecology 96:22032213.

Appendix C (Table C1). Parameters of SMS runs performed to estimate connectivity for the Cabanis’s greenbul and the natterjack toad.




Number of individuals released per site and per combination of parameter values



Method of estimation of effective cost value within the perceptual range

harmonic mean

harmonic mean

DP (directional persistence): tendency to follow a correlated path


2.0 to 16.0 in steps of 2.0

PR (perceptual range): extent of the landscape that can be perceived by the individual, and over which movement costs are estimated

25 m

30 m

Maximum number of steps allowed (per individual)

2 millions


Effect of landscape boundaries

totally reflective

partially absorbing

Specific biological adjustments

- implementation of a dispersal bias (=1.02, 1.04, 1.06, 1.08, 1.1, 1.14, 1.18), i.e., a tendency to orient the movement away from the natal location

- implementation of a memory size (=2), i.e., the number of previous steps over which the current direction is determined for the application of directional persistence

implementation of an “auditory attraction”: if the recruitment window is open (10000 steps / 100000), goal bias = 5.0 when an individual is within 1 km of a breeding site; recruitment not allowed when the recruitment window is closed; the goal bias is a tendency to orient the movement towards a particular location (in this case the center of the breeding site lying within the 1km auditory attraction buffer)

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