Ecological Archives E086-077-A3

Matthew A. Etterson and Richard S. Bennett. 2005. Including transition probabilities in nest survival estimation: a Mayfield Markov chain. Ecology 86:1414–1421.

Appendix C. Formulae for bias and standard errors of daily survival and overall success.

Parameter Quality Metric

Formula

Mean bias in parameter estimate

Mean standard error of daily survival

Mean standard error of overall success (delta method, e.g., Armstrong et al. 2002)

Standard deviation of parameter estimates

Formulae are identical for , with the estimate of interest substituted for  and the known expected value substituted for θ

Asymptotic standard error of is similarly estimated, but with the partial derivatives of  substituted for those of .



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