Ecological Archives E086-025-A7

James E. Byers. 2005. Marine reserves enhance abundance but not competitive impacts of a harvested nonindigenous species. Ecology 86:487–500.

Appendix G. A table of significant site-level covariates that were significant (or whose inclusion made a fixed factor significant) in exploratory analyses when added individually to the baseline mixed model (predator exposure and density) examining mortality and dry tissue growth responses of each species.

Protothaca mortality

Venerupis mortality

Protothaca growth

Venerupis growth

       

Predation × Organics

Predation × Crab biomass

Predation × Crab biomass

Predation × Crab numbers

Predation × Stdev Organics

Predation × Crab numbers

Predation × Crab numbers

Predation × Density × Crab numbers

Predation × Ln Venerupis Biomass

Predation × Organics

Predation × Stdev Chlorophyll

Predation × Ln Protothaca Biomass

Predation × Ln Protothaca Biomass

Predation × Stdev Organics

Predation × Ln Venerupis Biomass

Ln Protothaca Biomass

Ln Protothaca Biomass

Predation × Bird number

Predation × Density × Ln Venerupis Biomass

Predation × Ln Venerupis Biomass

Density × Ln Protothaca Biomass

Predation × Ln Venerupis Biomass

Predation × Total Clam Biomass

Density × Ln Protothaca Biomass

 

Crab biomass

Predation × Density × Total Clam Biomass

Predation × Density × Ln Venerupis Biomass

 

Bird number

Predation × Organics

 
 

Organics

Density × Ln Protothaca Biomass

 
   

Predation × Density × Ln Protothaca Biomass

 
   

Predation × Density × Exotic Biomass

 
   

Predation × Exotic Biomass

 

   Notes: Covariate terms are listed in order of significance. Factors significant to gonad growth were the same as those for growth for each species. Stdev is the standard deviation. The effect of crabs on both clam growth and mortality explains why the interaction of predator exposure and ambient Venerupis biomass significantly affected experimental clams. Notably, crab biomass is inversely related to Venerupis biomass (R2 = 0.46). Thus, when ambient Venerupis biomass was low, crabs were abundant, and both clams exhibited lower growth and survivorship in open topped enclosures that exposed them to predators.



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